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Note

Taxa Associated with the Family Gonatidae Hoyle, 1886

Michael J. Sweeney and Richard E. Young

(data as of June 2002)

Nominal Genus-Level Taxa

Berryteuthis Naef, 1921a:535.

Nomen nudum.

Berryteuthis Naef, 1921b:244.

CURRENT SYSTEMATIC STATUS. -- Valid genus [fide Nesis (1987:196)]
TYPE SPECIES. -- Gonatus magister Berry, 1913 by original designation

(Boreoteuthis) Nesis, 1971:65.

CURRENT SYSTEMATIC STATUS. -- Valid subgenus of Gonatopsis Sasaki, 1920 [fide Nesis (1987:196)]
TYPE SPECIES. -- Gonatopsis borealis Sasaki, 1923 by original designation

Chiloteuthis Verrill, 1881 In 1880-1881:292.

CURRENT SYSTEMATIC STATUS. -- Gonatus Gray, 1849 [as Cheloteuthis] [fide Hoyle (1910:408)]
TYPE SPECIES. -- Cheloteuthis rapax Verrill, 1881 by monotypy

(Eogonatus) Nesis, 1972:1300.

CURRENT SYSTEMATIC STATUS. -- Valid genus [fide Nesis (1987:197)]
TYPE SPECIES. -- Gonatus (Eogonatus) tinro Nesis, 1972 by monotypy

Gonatopsis Sasaki, 1920:198.

CURRENT SYSTEMATIC STATUS. -- Valid genus [fide Nesis (1987:196)]
TYPE SPECIES. -- Gonatopsis octopedatus Sasaki, 1920 by original designation

Gonatus Gray, 1849:67.

CURRENT SYSTEMATIC STATUS. -- Valid genus [fide Hoyle (1910:409)]
TYPE SPECIES. -- Onychoteuthis? amoena Moller, 1842 by monotypy

Lestoteuthis Verrill, 1880 In 1879-1880:251.

CURRENT SYSTEMATIC STATUS. -- Gonatus Gray, 1849 [fide Steenstrup (1882:146)]
TYPE SPECIES. -- Onychoteuthis kamschatica Middendorf, 1849 by original designation

Pfefferiopsis Ishikawa, 1924:185.

Nomen nudum; Berryteuthis Naef, 1921 [fide Ishikawa (1929:583)]
TYPE SPECIES. -- Gonatus magister Berry, 1913 by monotypy

 

Nominal Species-Level Taxa (As Introduced Binomial)

Onychoteuthis amoena Moller, 1842:76.

CURRENT SYSTEMATIC STATUS. -- Gonatus fabricii (Lichtenstein, 1818) [fide Morch (1857:89)]
TYPE REPOSITORY. -- BMNH Syntypes [3] 1843.7.3.74-6 [fide Lipinski et al. (2000:105)]
TYPE LOCALITY. -- West Greenland [fide Schiotte and Waren (1992:7)]

Gonatus anonychus Pearcy and Voss, 1963:105, figs 1-2.

CURRENT SYSTEMATIC STATUS. -- Valid species; Berryteuthis anonychus [fide Nesis (1987:197)]
TYPE REPOSITORY. -- NMNH Holotype 575559 [fide Roper and Sweeney (1978:4)]
TYPE LOCALITY. -- 45°26.8'N, 125°08.1'W (Pacific Ocean)

Gonatus antarcticus Lonnberg, 1898:51, pl 5, figs 4-5.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Nesis (1987:291)]
TYPE REPOSITORY. -- Unresolved; NRM? Holotype
TYPE LOCALITY. -- On beach, Punta Arenas, Chile

Gonatus berryi Naef, 1923:245.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Young (1972:46)]
TYPE REPOSITORY. -- NMNH Syntypes 214652 and 214658; specimens not extant
TYPE LOCALITY. -- Monterey Bay, California [fide Young (1972:48)]

Gonatopsis borealis Sasaki, 1923:203, text-fig.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Okutani et al. (1987:129)]
TYPE REPOSITORY. -- Unresolved; FMHU Syntypes[?]; [see Taki and Igarashi (1967:16)]
TYPE LOCALITY. -- Kushiro, Nemuro, Tokachi; all on eastern Hokkaido, Japan

Gonatus californiensis Young, 1972:51, pls 15b, 16b, 16j-p, 17d-f.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Nesis (1987:200)]
TYPE REPOSITORY. -- SBMNH Holotype 34978 [fide Scott et al. (1990:19)]
TYPE LOCALITY. -- 33°32'N, 118°24'W, eastern North Pacific Ocean

Onychoteuthis fabricii Lichtenstein, 1818:1592.

CURRENT SYSTEMATIC STATUS. -- Valid species; Gonatus fabricii [fide Berry (1912:308)]
TYPE REPOSITORY. -- ZMUC Neotype [fide Kristensen (1981:66)]
TYPE LOCALITY. -- Amerdloq Fjord, West Greenland [fide Kristensen and Knudsen (1983:222)]

Gonatopsis japonica Okiyama, 1969:20, text-figs 1-4, pls 1-3.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Nesis (1987:201)]
TYPE REPOSITORY. -- NSMT Holotype 60020 [fide Kubodera and Tsuchiya (1993:unpaginated)]
TYPE LOCALITY. -- 41°02'N, 138°11'E (Pacific Ocean)

Onychoteuthis kamtschatica Middendorff, 1849:515, pl 12.

CURRENT SYSTEMATIC STATUS. -- Valid species; Gonatus kamtschaticus [fide Nesis (1987:200)]
TYPE REPOSITORY. -- Unresolved
TYPE LOCALITY. -- Schumschu Island, Kurile Islands

Gonatus madokai Kubodera and Okutani, 1977:124, text-fig 1, pls 1-4.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Okutani et al. (1987:127)]
TYPE REPOSITORY. -- FMHU Holotype M-1362
TYPE LOCALITY. -- 49°32'N, 168°50'E (Northwest Pacific Ocean)

Gonatus magister Berry, 1913:76.

CURRENT SYSTEMATIC STATUS. -- Valid species; Berryteuthis magister [fide Naef (1921a:535)]
TYPE REPOSITORY. -- CASIZ Holotype 017965 [fide Sweeney et al. (1988:15)]
TYPE LOCALITY. -- Puget Sound, Washington

Gonatopsis borealis makko Okutani and Nemoto, 1964:113, pl 4.

CURRENT SYSTEMATIC STATUS. -- Valid species; Gonatopsis makko [fide Nesis (1987:201)]
TYPE REPOSITORY. -- NSMT Holotype Mo-13311
TYPE LOCALITY. -- Southern Bering Sea (from sperm whale stomach)

Gonatus middendorffi Kubodera and Okutani, 1981:8, pl 1.

CURRENT SYSTEMATIC STATUS. -- Gonatus kamtschaticus (Middendorff, 1849) [fide Nesis (1987:200)]
TYPE REPOSITORY. -- NSMT Holotype Mo-58621 [fide Kubodera and Tsuchiya (1993:unpaginated)]
TYPE LOCALITY. -- 45°30'N, 151°40'E (Pacific Ocean)

Berryteuthis magister nipponensis Okutani and Kubodera, 1987:133, figs a-c.

CURRENT SYSTEMATIC STATUS. -- Valid subspecies [fide; taxon not yet reviewed]
TYPE REPOSITORY. -- NSMT Holotype 64264 [fide Kubodera and Tsuchiya (1993:unpaginated)]
TYPE LOCALITY. -- Okirai Bay, Iwate, northeastern Honshu, Japan

Gonatopsis octopedatus Sasaki, 1920:198, pl 26 fig 5.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Nesis (1987:201)]
TYPE REPOSITORY. -- NMNH Holotype 332918 [fide Roper and Sweeney (1978:10)]
TYPE LOCALITY. -- Near Cape Patience, Sakhalin (48°22'30"N, 145°43'30"E), Okhotsk Sea

Gonatopsis (Gonatopsis) okutanii Nesis, 1972:1304, fig 2.

CURRENT SYSTEMATIC STATUS. -- Valid species? [fide Nesis (1987:201)]
TYPE REPOSITORY. -- ZIN Holotype
TYPE LOCALITY. -- 44°06.4'N, 150°28.2'E (Pacific Ocean)

Gonatus onyx Young, 1972:43, pls 13a, 14a-c, 14i, 17j.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Okutani et al. (1987:125)]
TYPE REPOSITORY. -- SBMNH Holotype 34969 [fide Scott et al. (1990:19)]
TYPE LOCALITY. -- 33°19'N, 118°45'W, eastern North Pacific Ocean

Gonatus oregonensis Jefferts, 1985:167, figs 4-5.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide; taxon not yet reviewed]
TYPE REPOSITORY. -- CASIZ Holotype 040162
TYPE LOCALITY. -- Off Oregon, U.S.A.

Gonatus phoebetriae Imber, 1978:480, fig 7c.

CURRENT SYSTEMATIC STATUS. -- Species dubium [fide Jefferts (1985:159)]
TYPE REPOSITORY. -- Percy Fitzpatrick Institute Holotype (single lower beak)
TYPE LOCALITY. -- Marion Is., 46°55'S, 37°45'E (S Indian Ocean) (from stomach of sooty albatross, Phoebetria fusca)

Gonatus pyros Young, 1972:49, pls 13b, 14b, 14j-q, 17g-i, 17k.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Nesis (1987:200)]
TYPE REPOSITORY. -- SBMNH Holotype 34973 [fide Scott et al. (1990:19)]
TYPE LOCALITY. -- 33°37'N, 118°26'W, eastern North Pacific Ocean

Cheloteuthis rapax Verrill, 1881 In 1880-1881:293, pl 49 fig 1-1f.

CURRENT SYSTEMATIC STATUS. -- Gonatus fabricii (Lichtenstein, 1818) [fide Berry (1912:308)]
TYPE REPOSITORY. -- NMNH Holotype 574639 [fide Roper and Sweeney (1978:12)]
TYPE LOCALITY. -- 39°52'20"N, 70°58'00"W (Atlantic Ocean)

Gonatus septemdentatus Sasaki, 1915:185.

CURRENT SYSTEMATIC STATUS. -- Berryteuthis magister (Berry, 1913) [fide Nesis (1987:196)]
TYPE REPOSITORY. -- Unresolved; Syntypes (6)
TYPE LOCALITY. -- Toyama Bay, Etchu Province, Japan

Berryteuthis magister shevtsovi Katugin, 2000:91, figs 7-11.

CURRENT SYSTEMATIC STATUS. -- Valid subspecies [fide; taxon not yet reviewed]
TYPE REPOSITORY. -- TINRO Holotype BMJS001
TYPE LOCALITY. -- Japan Sea, 42°30'N, 113°42'E

Gonatus steenstrupi Kristensen, 1981:78, figs 18-28.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Nesis (1987:200)]
TYPE REPOSITORY. -- ZMUC Holotype [fide Kristensen and Knudsen (1983:224)]
TYPE LOCALITY. -- Between 53°24' - 51°39'N and 15°52' - 8°29'W (Northeast Atlantic Ocean)

Gonatus (Eogonatus) tinro Nesis, 1972:1300, fig 1.

CURRENT SYSTEMATIC STATUS. -- Valid species; [fide Nesis (1987:197)]
TYPE REPOSITORY. -- ZIN Holotype
TYPE LOCALITY. -- Bering Sea, 58°22.2'N, 174°55'W

Gonatus ursabrunae Jefferts, 1985:160, figs 1-2.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide; taxon not yet reviewed]
TYPE REPOSITORY. -- CASIZ Holotype 040163
TYPE LOCALITY. -- South of Alaska Peninsula

 

Type Repositories

BMNH-The Natural History Museum, Cromwell Road, London SW7 5BD, England (formerly British Museum (Natural History)).† [For type catalog see Lipinski et al. (2000)]

CASIZ-California Academy of Sciences, Golden Gate Park, San Francisco, CA 94118, U.S.A.† [Smith (1974); Sweeney et al. (1988) for S.S. Berry taxa]

FMHU- Fisheries Museum, Faculty of Fisheries, Hokkaido University, Nishi 5, Kita 8, Sapporo 060, Hakodate, Japan (includes Laboratory of Marine Zoology (HUMZ) collection).† [For type catalog see Taki and Igarashi (1967)]

NMNH-National Museum of Natural History, 10th & Constitution Avenue, Smithsonian Institution, Washington, DC 20560, U.S.A. (formerly USNM).† [For type catalogs see Roper and Sweeney (1978); Sweeney et al. (1988)]

NRM- Naturhistoriska Riksmuseet, Box 50007, S-104 05 Stockholm, Sweden. [For type catalog of Odhner taxa see Sandberg and Waren (1993)]

NSMT- National Science Museum, 3-23-1 Hyakunin-cho, Shinjuku-ku, Tokyo 160, Japan.† [For type catalog see Kubodera and Tsuchiya (1993)]

SBMNH-Santa Barbara Museum of Natural History, 2559 Puesta del Sol Road, Santa Barbara, CA 93105, U.S.A.† [For type catalogs see Hertz (1984); Scott et al. (1990)]

TINRO- Pacific Research Fisheries Center, Vladivostok, Russia.

ZIN- Zoological Institute, Russian Academy of Sciences, Universitetskaya Nab. 1, St. Petersburg 199164, Russia.

ZMUC-Kobenhavns Universitet, Zoologisk Museum, Universitetsparken 15, DK 2100 Copenhagen, Denmark.† [For type catalog see Kristensen and Knudsen (1983); See also Volsoe et al. (1962) for Steenstrup papers]

References

Berry, S.S. 1912. A review of the cephalopods of western North America. Bulletin of the Bureau of Fisheries, 30(1910):267-336, 18 figures, 25 plates.

Berry, S.S. 1913. Notes on some West American cephalopods. Proceedings of the Academy of Natural Sciences of Philadelphia, 65:72-77, 2 figures.

Gray, J.E. 1849. Catalogue of the Mollusca in the British Museum. Part I. Cephalopoda Antepedia. 164 pages. London.

Hertz, C.M. 1984. Illustrations of the types named by S. Stillman Berry in his “Leaflets in Malacology.” Festivus, 15 (Suppl.):1-42.

Hoyle, W.E. 1886. Report on the Cephalopoda. Report on the scientific results of the voyage of H.M.S. Challenger during the years 1873-76, Zoology, 16(44):1-245, 33 plates.

Hoyle, W.E. 1910. A list of the generic names of Dibranchiate Cephalopoda with their type species. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 32:407-413.

Imber, M.J. 1978. The squid families Cranchiidae and Gonatidae (Cephalopoda: Teuthoidea) in the New Zealand region. New Zealand Journal of Zoology, 5:445-484, 7 figures.

Ishikawa, M. 1924. On the phylogenetic position of the cephalopod genera of Japan based on the structure of statocysts. Journal of the College of Agriculture, Imperial University of Tokyo, 7(3):165-210, 6 plates.

Ishikawa, M. 1929. On the statocyst of the American cephalopod genera. Journal of Morphology and Physiology, 48(2):563-584.

Jefferts, K. 1985. Gonatus ursabrunae and Gonatus oregonensis, two new species from the northeastern Pacific Ocean (Cephalopoda: Oegopsida: Gonatidae). Veliger, 28(2):159-174, 5 figures.

Katugin, O.N. 2000. A new species of the schoolmaster gonate squid, Berryteuthis magister (Cephalopoda: Gonatidae), from the Japan Sea. The Veliger, 43(1):82-97, 11 figures.

Kristensen, T.K. 1981. The genus Gonatus Gray, 1849 (Mollusaca: Cephalopoda) in the North Atlantic. A revision of the North Atlantic species and description of Gonatus steenstrupi n. sp. Steenstrupia, 7(4):61-99, 29 figures.

Kristensen, T.K. and J. Knudsen. 1983. A catalogue of the type specimens of Cephalopoda (Mollusca) in the Zoological Museum, University of Copenhagen. Steenstrupia, 9(10):217-227.

Kubodera, T. and T. Okutani. 1977. Description of a new species of gonatid squid, Gonatus madokai, n. sp., from the northwest Pacific, with notes on morphological changes with growth and distribution in immature stages (Cephalopoda: Oegopsida). Venus, 36(3):123-151, 3 figures, 4 plates.

Kubodera, T. and T. Okutani. 1981. Gonatus middendorffi, a new species of gontid squid from the northern North Pacific, with notes on morphological changes with growth and distribution in immature stages (Cephalopoda: Oegopsida). Bulletin of the National Science Museum, Tokyo, (series A)7(1):7-26, 1 plate.

Kubodera, T., and K. Tsuchiya (eds.) 1993. Catalogue of Specimens of the Class Cephalopoda (Phylum Mollusca) in the National Science Museum, Tokyo. National Science Museum, Tokyo, 1993, unpaginated.

Lichtenstein, H.C. 1818. Onychoteuthis, Sepien mit Krallen. Isis oder Encyclopadische Zeitung, 1818(9):1591-1592, 1 plate.

Lipinski, M.R., F.A. Naggs, and M.A. Roeleveld. 2000. Catalogue of types of Recent cephalopods in the collection of the Natural History Museum, London. Annales Zoologici (Warszawa), 50(1):101-120.

Lönnberg, E. 1898. On the cephalopods collected by the Swedish Expedition to Tierra del Fuego, 1895-96. Svenska Expeditionen till Magellanslanderna, 2(4):49-64, 2 plates.

Middendorff, A.T. 1849. Beitrage zu einer Malacozoologica Rossica. II-III. Aufzahlung und Beschreibung der zur Meeresfauna Russlands gehorigen Einschaler. Memoires de l'Academie imperiale des Sciences de St. Petersbourg, (series 6)6:329-610, 21 plates.

Moller, H.P.C. 1842. Index Molluscorum Groenlandiae. Naturhistorisk Tidskrift, Copehagen, 4(1):76-97.

Morch, O.A.L. 1857. Fortegnelse over Gronlands bloddyr. Tilloeg nr. 4, pages 75-100. In: Rink, Gronland Geographisk og Statistisk beskrivet af Rink.

Naef, A. 1921a. Das system der dibranchiaten Cephalopoden und die mediterranen arten derselben. Mitteilungen aus der Zoologischen Station zu Neapel, 22(16):527-542, 1 figure.

Naef, A. 1921b. Die Cephalopoden. Fauna und Flora des Golfes von Neapel, Monograph 35, 1(1)(part 1):1-148, figures 1-62, plates 1-56.

Naef, A. 1923. Die Cephalopoden. Fauna e Flora de Golfo di Napoli, Monograph 35, 1(1)(part 2):149-863, figures 63-473.

Nesis, K.N. 1971. The family Gonatidae - abundant squids of the North Pacific (their distribution, ecology, systematics and phylogeny), pages 63-65. Fourth Meeting of the Investigation of Molluscs: Molluscs trends, methods and some results of their distribution. Moscow: Academy of Sciences USSR Zoological Institute.

Nesis, K.N. 1972. Two new species of gonatid squids from the North Pacific. Zoologichesky Zhurnal, 51(9):1300-1307, 2 figures. [In Russian]

Nesis, K.N. 1982. Abridged key to the cephalopod Molluscs of the world's oceans. Moscow: Light and Food Industry Publishing House, 385 pages. (In Russian; see Nesis, 1987 for abridged English translation)

Nesis, K.N. 1987. Cephalopods of the World; Squids, cuttlefishes, Octopuses, and Allies. T.F.H. Publications, Neptune City, NJ, USA, 351 pages.

Okiyama, M. 1969. A new species of Gonatopsis from the Japan Sea, with the record of a specimen referable to Gonatopsis sp. Okutani 1967 (Cephalopoda: Oegopsida: Gonatidae). Publications of the Seto Matine Biological Laboratory, 17(1):19-32, 4 figures, 4 plates.

Okutani, T. and T. Kubodera. 1987. In: T. Okutani, M. Tagawa and H. Horikawa, Cephalopods from continental shelf and slope around Japan. 194 pages. Tokyo: Japan Fisheries Resource Conservation Association.

Okutani, T. and T. Nemoto. 1964. Squids as the food of sperm whales in the Bering Sea and Alaskan Gulf. Scientific Reports of the Whales Research Institute, 18:111-121, 5 plates.

Okutani, T., M. Tagawa, and H. Horikawa. 1987. Cephalopods from continental shelf and slope around Japan. 194 pages. Tokyo: Japan Fisheries Resource Conservation Association.

Pearcy, W.G. and G.L. Voss. 1963. A new species of gonatid squid from the northeastern Pacific. Proceedings of the Biological Society of Washington, 76:105-112, 2 figures.

Roper, C.F.E. and M.J. Sweeney. 1978. A catalog of the type-specimens of Recent Cephalopoda in the National Museum of Natural History. Smithsonian Contributions to Zoology, 278:1-19.

Sandberg, L. and A. Waren. 1993. Molluscan taxa introduced by Nils Hjalmar Odhner. Malakologische Abhandlungen Staatliches Museum fur Tierkkunde Dresden, 16(15):117-134.

Sasaki, M. 1915. On a new species of oegopsid from the Bay of Toyama, Gonatus septemdentatus. Transactions of the Sapporo Natural History Society, 5(3):185-189.

Sasaki, M. 1920. Report on cephalopods collected during 1906 by the United States Bureau of Fisheries steamer "Albatross" in the northwestern Pacific. Proceedings of the United States National Museum, 57(2310):163-203, 4 plates.

Sasaki, M. 1923. On a new eight-armed squid from Hokkaido, Gonatopsis borealis n. sp. Annotationes Zoologicae Japonenses, 10:203-207, 1 figure.

Schiotte, T. and A. Waren. 1992. An annotated and illustrated list of the types of Mollusca described by H.P.C. Moller from West Greenland. Meddelelser om Gronland, Bioscience, 35:1-34, 119 figures.

Scott, P.H., F.G. Hochberg and B. Roth. 1990. Catalog of Recent and fossil molluscan types in the Santa Barbara Museum of Natural History. I. Caudofoveata, Polyplacophora, Bivalvia, Scaphopoda and Cephalopoda. The Veliger, 33(Suppl. 1):1-27.

Smith, A.G. 1974. Type and other cephalopods in the collection of the Department of Invertebrate Zoology, California Academy of Sciences. Veliger, 16(3):319-333.

Steenstrup, J. 1882. Notae teuthologicae, 3. Oversigt over det Kongelige Danske Videnskabernes Selskabs Forhandlinger, 1882:53-164.

Sweeney, M.J., C.F.E. Roper, and F.G. Hochberg. 1988. Catalog of the type specimens of Recent Cephalopoda described by S. Stillman Berry. Malacologia, 29(1):7-19.

Taki, I. and T. Igarashi. 1967. A list of cephalopod specimens in the Fisheries Museum, Faculty of Fisheries, Hokkaido University (Sasaki collection and specimens collected by the Marine Zoological Laboratory). Fisheries Museum, Faculty of Fisheries, Hokkaido University, Contribution, 7:1-27.

Verrill, A.E. 1879-1880. The cephalopods of the north-eastern coast of America. Part 1. - The gigantic squids (Architeuthis) and their allies; with observations on similar large species from foreign localities. Transactions of the Connecticut Academy of Sciences, 5(5):177-257, 14 plates.

Verrill, A.E. 1880-1881. The cephalopods of the north-eastern coast of America. Part II. The smaller cephalopods, including the "squids" and the octopi, with other allied forms. Transactions of the Connecticut Academy of Sciences, 5(6):259-446, 33 plates.

Volsoe, A., J. Knudsen and W. Rees. 1962. The cephalopod papers of Japetus Steenstrup; a translation into English. Danish Science Press, Copenhagen, 330 pages.

Young, R.E. 1972. The systematics and areal distribution of pelagic cephalopods from the seas off southern California. Smithsonian Contributions to Zoology, 97:1-159, 15 figures, 38 plates.

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